In the aftermath of the plane crash, the residents of Wisteria Lane think about what their lives might have been like had they made different choices.
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Susan turns up the heat for Mike, Bree learns it won’t be easy to undo the hurt she’s caused Orson, Lynette discovers Tom’s true intentions, and Gaby is forced to examine feelings she’s suppressed about her heritage.
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On the highly anticipated season five premiere, five years have passed since Katherine shot her ex-husband as Susan, Lynette, Bree and Gabrielle stood together in solidarity to protect her. Susan tries to hide her relationship with a new man. Gaby questions her abilities as a mother. Bree is on the verge of publishing her own cookbook as business partner Katherine grows envious. Lynette catches her teenaged twins spearheading illegal activities. And finally, Edie returns to Wisteria Lane in jaw-dropping fashion with a new man in tow, her new husband Dave (played by Neal McDonough).
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Lynette discovers her son, Porter, has a softer side, which she finds endearing. Gaby amusingly attempts to mask her demise in social stature. The time has come for the men in Susan’s life to meet. Meanwhile, Bree begins to feel the pressures of balancing marriage and a career. Edie is reminded just how hostile the neighbors on Wisteria Lane can be. Finally, a suspicious Mrs. McCluskey enlists Katherine’s help to uncover more about their new neighbor, Dave.
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Bree’s estranged daughter, Danielle, returns to Wisteria Lane and is met with a deluge of criticism. Meanwhile, Gabrielle and Susan find themselves embroiled in a catfight brought on by an altercation between their kids. To her chagrin, Lynette comes home to find Tom and Dave have formed a garage band after she told him to get rid of his old things. Finally, Katherine and Mrs. McCluskey invite Edie to lunch to press her for more information on Dave.
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Bree’s new found success causes the rest of the women on Wisteria Lane to feel envious. Lynette longs to return to her days as an ad-executive, and finds an outlet as Bree’s marketing manager. Mike feels that he is not a big enough presence in his son’s life after M.J. draws a family portrait and leaves him out. Meanwhile, Gaby comes up with a way to have more sex with Carlos.
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Driven by a hidden agenda, Dave convinces Susan to throw Mrs. McCluskey a surprise party for her 70th birthday. Jackson wants to take his relationship with Susan to the next level. Gabrielle’s concerned she might be pregnant. Tom tells Lynette he’s sold the pizzeria. A startling reason for the bond between Bree and Katherine is revealed. And finally, an outraged Mrs. McCluskey has the entire neighborhood questioning her sanity.
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Gaby is over the moon when Carlos’ client makes them an offer they can’t refuse, but a startling discovery leaves her second guessing. Lynette grows suspicious that Tom is cheating on her. After breaking up with Jackson, Susan tries to surprise him, only to find the surprise is on her. Meanwhile, the bond between Bree and Katherine grows. Finally, Mrs. McCluskey enlists the help of her sister Roberta (guest star Lily Tomlin) to dig up dirt on Dave.
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Lynette and Tom uncover the unsettling truth about son Porter. Gabrielle suspects Carlos’ best client, Virginia (guest star Frances Conroy), has ulterior motives. Bree’s moment of weakness could bring her great embarrassment and shame. Katherine’s not-so-little secret is unveiled. Susan learns of Jackson’s true passion. Finally, Mrs. McCluskey’s sister Roberta (guest star Lily Tomlin) unearths a disturbing fact about Dave.
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There’s a big fire at a club where the men’s band is playing in a battle of the bands. One of the other bands competing is real-life band Tokyo Police Club. Most residents of Wisteria Lane will be present.
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